John R. Anderson Member (elected 1999)
Carnegie Mellon University
Primary: 52, Psychological and Cognitive Sciences

Mary T. K. Arroyo International Member (elected 1999)
University of Chile
Primary: 63, Environmental Sciences and Ecology

Richard A. Askey Member (elected 1999)
June 4, 1933 - October 9, 2019

Robert H. Austin Member (elected 1999)
Princeton University
Primary: 13, Physics

Bernard M. Babior Member (elected 1999)
November 10, 1935 - June 29, 2004

William A. Bardeen Member (elected 1999)
Fermi National Accelerator Laboratory
Primary: 13, Physics

Marlene Belfort Member (elected 1999)
University at Albany, State University of New York
Primary: 21, Biochemistry
Secondary: 26, Genetics

Elwyn Berlekamp Member (elected 1999)
September 6, 1940 - April 9, 2019

Michael J. Berridge International Member (elected 1999)
October 22, 1938 - February 13, 2020

Robert L. Carneiro Emeritus (elected 1999)
June 4, 1927 - June 24, 2020

Joanne Chory Member (elected 1999)
Salk Institute for Biological Studies
Primary: 25, Plant Biology

James E. Cleaver Member (elected 1999)
University of California, San Francisco
Primary: 41, Medical Genetics, Hematology, and Oncology
Secondary: 26, Genetics

John M. Coffin Member (elected 1999)
Tufts University
Primary: 44, Microbial Biology
Secondary: 41, Medical Genetics, Hematology, and Oncology

William F. DeGrado Member (elected 1999)
University of California, San Francisco
Primary: 29, Biophysics and Computational Biology

Charles H. DePuy Member (elected 1999)
September 10, 1927 - March 14, 2013

Robert Desimone Member (elected 1999)
Massachusetts Institute of Technology
Primary: 28, Systems Neuroscience

Patricia K. Donahoe Member (elected 1999)
Harvard University
Primary: 42, Medical Physiology and Metabolism

Joseph Felsenstein Member (elected 1999)
University of Washington
Primary: 27, Evolutionary Biology
Secondary: 32, Applied Mathematical Sciences

Stephen E. Fienberg Member (elected 1999)
November 27, 1942 - December 14, 2016

Paul A. Fleury Member (elected 1999)
Yale University
Primary: 33, Applied Physical Sciences

Howard M. Grey Member (elected 1999)
August 16, 1932 - December 22, 2019

Richard S. Hamilton Member (elected 1999)
Columbia University
Primary: 11, Mathematics

Bruce D. Hammock Member (elected 1999)
University of California, Davis
Primary: 61, Animal, Nutritional, and Applied Microbial Sciences

Thomas J. Hanratty Emeritus (elected 1999)
November 9, 1926 - August 24, 2016

Lars Peter Hansen Member (elected 1999)
The University of Chicago
Primary: 54, Economic Sciences

Wick C. Haxton Member (elected 1999)
University of California, Berkeley
Primary: 13, Physics
Secondary: 12, Astronomy

Ralph F. Hirschmann Member (elected 1999)
May 6, 1922 - June 20, 2009

Albrecht W. Hofmann International Member (elected 1999)
Max Planck Institute for Chemistry
Primary: 15, Geology

Tim Hunt International Member (elected 1999)
The Francis Crick Institute
Primary: 22, Cellular and Developmental Biology
Secondary: 21, Biochemistry

Louis J. Ignarro Member (elected 1999)
University of California, Los Angeles
Primary: 23, Physiology and Pharmacology
Secondary: 21, Biochemistry

Vaughan F. Jones Member (elected 1999)
December 31, 1952 - September 6, 2020

C. Ronald Kahn Member (elected 1999)
Harvard University
Primary: 42, Medical Physiology and Metabolism
Secondary: 22, Cellular and Developmental Biology

Arthur Karlin Member (elected 1999)
Columbia University
Primary: 23, Physiology and Pharmacology

Margaret G. Kivelson Member (elected 1999)
University of California, Los Angeles
Primary: 16, Geophysics

Arthur Landy Member (elected 1999)
Brown University
Primary: 21, Biochemistry
Secondary: 26, Genetics

Ramon Latorre International Member (elected 1999)
Universidad de Valparaiso
Primary: 23, Physiology and Pharmacology
Secondary: 29, Biophysics and Computational Biology

Raphael D. Levine International Member (elected 1999)
The Hebrew University of Jerusalem
Primary: 14, Chemistry

Douglas K. Lilly Member (elected 1999)
June 16, 1929 - June 24, 2018

Steven E. Lindow Member (elected 1999)
University of California, Berkeley
Primary: 62, Plant, Soil, and Microbial Sciences

Akinlawon Mabogunje International Member (elected 1999)
October 18, 1931 - August 4, 2022

Enid MacRobbie International Member (elected 1999)
University of Cambridge
Primary: 25, Plant Biology

Dusa McDuff Member (elected 1999)
Barnard College
Primary: 11, Mathematics

J. Richard McIntosh Member (elected 1999)
University of Colorado Boulder
Primary: 22, Cellular and Developmental Biology
Secondary: 29, Biophysics and Computational Biology

Michael M. Merzenich Member (elected 1999)
Posit Science Corporation
Primary: 28, Systems Neuroscience

Janice M. Miller Member (elected 1999)
U.S. Department of Agriculture
Primary: 61, Animal, Nutritional, and Applied Microbial Sciences

James A. Mirrlees International Member (elected 1999)
July 5, 1936 - August 29, 2018

Cherry A. Murray Member (elected 1999)
University of Arizona
Primary: 33, Applied Physical Sciences
Secondary: 31, Engineering Sciences

Satoshi Omura International Member (elected 1999)
Kitasato University
Primary: 61, Animal, Nutritional, and Applied Microbial Sciences
Secondary: 14, Chemistry

Thomas D. Petes Member (elected 1999)
Duke University
Primary: 26, Genetics
Secondary: 44, Microbial Biology

Michael E. Phelps Member (elected 1999)
University of California, Los Angeles
Primary: 31, Engineering Sciences